Hypopitys lanuginosa

Family:ERICACEAE
Species:Hypopitys lanuginosa (Michx.) Raf.
Common Name:PINESAP; FALSE BEECHDROPS
Plant Notes:This achlorophyllous mycoheterotrophic species was found to most often associate with ectomycorrhizal Tricholoma (Bidartondo & Bruns 2001; Johansson et al. 2017). This species is now put in the genus Hypopitys which is more closely related to Pityopus than Monotropa (Bidartondo & Bruns 2001). Tentatively, the Florida representative is here considered H. lanuginosa based on the study by Broe (2014), however the extensive synonymy list of the previously recognized Hypopitys monotropa s.l. is here retained until the taxonomy is thoroughly revised. Also see the treatment by Small (1914).
Status:Native, Endangered-State
Specimen: View specimen details in the USF Herbarium Specimen Database

** Not applicable or data not available.

Classification

Division
Class
Sub Class
Order ERICALES
FamilyERICACEAE
Genus Hypopitys
Species Hypopitys lanuginosa (Michx.) Raf. - PINESAP; FALSE BEECHDROPS

Citation

Citation Hypopitys lanuginosa (Michaux) Rafinesque, Med. Repos., ser. 3. 1: 297. 1810.
Basionym: Monotropa lanuginosa Michaux 1803.
Type: NORTH CAROLINA: "sylvis" (holotype: P [P00650948]).

** Not applicable or data not available.

Source

CountyHerbariaLiterature Citation (If Applicable)
Lake USF  
Marion USF  
Okaloosa USF  
Walton USF  

Synonyms

– Denotes synonyms that are applicable to the state. Show these synonyms only

SynonymFull CitationBasionymType
Hypopitys europaea Hypopitys europaea Nuttall, Gen. N. Amer. Pl. 1: 271. 1818, nom. illegit.BASIONYM: Hypopitys monotropa Crantz 1766; Monotropa hypopithys Linnaeus 1753. 
Hypopitys hypopithys Hypopitys hypopithys (Linnaeus) Small, Mem. Torrey Bot. Club 4: 137. 1893, nom. inadmiss.BASIONYM: Monotropa hypopithys Linnaeus 1753. 
Hypopitys lutea Hypopitys lutea Gray, Nat. Arr. Brit. Pl. 2: 404. 1821, nom. illegit.BASIONYM: Hypopitys monotropa Crantz 1766; Monotropa hypopithys Linnaeus 1753. 
Hypopitys monotropa HYPOPITYS MONOTROPA Crantz, Inst. Rei. Herb. 2: 467. 1766.Monotropa hypopithys Linnaeus 1753.Without data, Herb. Burser III: 156 (lectotype: UPS). Lectotypified by G. D. Wallace, in Luteyn, Fl. Neotrop. 66: 20. 1995.
Hypopitys multiflora Hypopitys multiflora Scopoli, Fl. Carniol., ed. 2. 1: 285. 1771, nom. illegit.BASIONYM: Hypopitys monotropa Crantz 1766; Monotropa hypopithys Linnaeus 1753. 
Hypopitys multiflora var. hirsuta Hypopitys multiflora Scopoli, var. hirsuta Ledebour, Fl. Ross. 2: 934. 1846, nom. inadmiss.BASIONYM: Monotropa hypopithys Linnaeus, var. hirsuta Roth 1788, nom. inadmiss.; Monotropa hypopithys Linnaeus 1753. 
Monotropa europaea Monotropa europaea Eaton & J. Wright, Man. Bot., ed. 8. 323. 1840, nom. illegit.BASIONYM: Hypopitys europaea Nuttall 1818, nom. illegit.; Hypopitys monotropa Crantz 1766; Monotropa hypopithys Linnaeus 1753. 
Monotropa hirsuta Monotropa hirsuta Hornemann, Fors. Oecon. Plantel., ed. 3. 2: 179. 1835, nom. illegit.BASIONYM: Monotropa hypopithys Linnaeus, var. hirsuta Roth 1788, nom. inadmiss.; Monotropa hypopithys Linnaeus 1753. 
Monotropa hypopithys Monotropa hypopithysLinnaeus, Sp. Pl. 387. 1753.  
Monotropa hypopithys subsp. lanuginosa Monotropa hypopithys Linnaeus, subsp. lanuginosa (Michaux) H. Hara, J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 6: 348. 1956.BASIONYM: Monotropa lanuginosa Michaux 1803. 
Monotropa hypopithys var. hirsuta Monotropa hypopithys Linnaeus, var. hirsuta Roth, Tent. Fl. Germ. 1: 180. 1788, nom. inadmiss.BASIONYM: Monotropa hypopithys Linnaeus 1753. 
Monotropa lanuginosa Monotropa lanuginosa Michaux, Fl. Bor.-Amer. 1: 266. 1803. TYPE: NORTH CAROLINA:
Monotropa multiflora Monotropa multiflora Fritsch, Excursionsfl. Oesterreich 426. 1897, nom. illegit.BASIONYM: Hypopitys multiflora Scopoli 1771, nom. illegit.; Hypopitys monotropa Crantz 1766; Monotropa hypopithys Linnaeus 1753. 
Monotropa squamiformis Monotropa squamiformis Dulac, Fl. Hautes-Pyrénées 421. 1867.BASIONYM: Monotropa hypopithys Linnaeus 1753. 
Hypopitys americana Hypopitys americana (de Candolle) Small, Fl. S.E. U.S. 880, 1336. 1903.BASIONYM: Hypopitys multiflora Scopoli, var. americana de Candolle 1839. 
Hypopitys brevis Hypopitys brevis Small, in Britton, N. Amer. Fl. 29(1): 13. 1914. TYPE: OREGON: Marion Co.: Independence Valley, Aug 1892, Craig 5214 (holotype: NY; isotype: OSC).
Hypopitys dentata Hypopitys dentata Rafinesque, Med. Repos., ser. 3. 1: 297. 1810. TYPE: "Virginia and Maryland",
Hypopitys fimbriata Hypopitys fimbriata (A. Gray) Howell, Fl. N.W. Amer. 429. 1901.BASIONYM: Monotropa fimbriata A. Gray 1873. 
Hypopitys glabra Hypopitys glabra (Roth) de Candolle, Prodr. 7: 780. 1839, nom. illegit.BASIONYM: Monotropa hypophegea Wallroth 1822; Monotropa hypopithys Linnaeus, var. glabra Roth 1788. 
Hypopitys hypophegea Hypopitys hypophegea (Wallroth) G. Don, Gen. Hist. 3: 866. 1834.BASIONYM: Monotropa hypophegea Wallroth 1822; Monotropa hypopithys Linnaeus, var. glabra Roth 1788. 
Hypopitys insignata Hypopitys insignata E. P. Bicknell, Bull. Torrey Bot. Club 41: 413. 1914. TYPE: MASSACHUSETTS: Dukes Co.: Chappaquiddick Island, 5 Oct 1912, Bicknell 6775 (holotype: NY; isotypes: NY 2 shts).
Hypopitys lanuginosa var. rosea Hypopitys lanuginosa (Michaux) Small, var. rosea House, Bull. New York State Mus. 243/244: 69. 1923, nom. illegit.BASIONYM: Monotropa lanuginosa Michaux, var. rubra Torrey 1846. 
Hypopitys latisquama Hypopitys latisquama Rydb., Bull. Torrey Bot. Club 41: 461. 1913. TYPE: MONTANA: Gallatin Co.: Bridger Mountains, 28 Jul 1896, Flodman 708 (holotype: NY; isotype: NY).
Hypopitys monotropa var. glabra Hypopitys monotropa Crantz, var. glabra (Roth) Druce, List. Brit. Pl. 47. 1908.BASIONYM: Monotropa hypopithys Linnaeus, var. glabra Roth 1788. 
Hypopitys multiflor var. serotina Hypopitys multiflora Scopoli, var. serotina Rouy, in Rouy & Foucaud, Fl. France 4: 10. 1897.  
Hypopitys multiflora subsp. hypophegea Hypopitys multiflora Scopoli, subsp. hypophegea (Wallroth) Rouy, in Rouy & Foucaud, Fl. France 4: 10. 1897.BASIONYM: Monotropa hypophegea Wallroth 1822; Monotropa hypopithys Linnaeus, var. glabra Roth 1788. 
Hypopitys multiflora var. americana Hypopitys multiflora Scopoli, var. americana de Candolle, Prodr. 7: 780. 1839.  
Hypopitys multiflora var. glabra Hypopitys multiflora Scopoli, var. glabra (Roth) Ledebour, Fl. Ross. 2: 934. 1846.BASIONYM: Monotropa hypopithys Linnaeus, var. glabra Roth 1788. 
Hypopitys multiflora var. ramosa Hypopitys multiflora Scopoli, var. ramosa Rouy, in Rouy & Foucaud, Fl. France 4: 10. 1897.  
Hypopitys racemosa Hypopitys racemosa Rafinesque, Med. Repos., ser. 3. 1: 297. 1810. TYPE: "Maryland, Pennsylvania",
Hypopitys rivinii Hypopitys rivinii Ruppius f. ex Cesati et al., Comp. Fl. Ital., ed. 1886. 418. 1886.  
Hypopitys secunda Hypopitys secunda Rafinesque, Med. Repos., ser. 3. 1: 297. 1810. TYPE: "Carolina",
Hypopitys stricta Hypopitys stricta Rafinesque, Med. Repos., ser. 3. 1: 297. 1810. TYPE: MARYLAND:
Monotropa abietina Monotropa abietina Dumortier, Bull. Soc. Roy. Bot. Belgique 4: 342. 1865.  
Monotropa abietina var. sanguinea Monotropa abietina Dumortier, var. sanguinea Haussknecht, Mitt. Geogr. Ges. (Thueringen) Jena 3: 286. 1885.  
Monotropa chinensis Monotropa chinensis Koidzumi, Fl. Symb. Orient.-Asiat. 28. 1930. TYPE: CHINA:
Monotropa fimbriata Monotropa fimbriata A. Gray, Proc. Amer. Acad. Arts 8: 629. 1873. TYPE: OREGON: Marion Co.: Silverton, 30 Jun 1871, Hall 357 (holotype: GH; isotype: NY).
Monotropa glabra Monotropa glabra (Roth) Bernhardi ex Reichenbach, Fl. Germ. Excurs. 411. 1831-1832, nom. illegit.BASIONYM: Monotropa hypophegea Wallroth 1822; Monotropa hypopithys Linnaeus, var. glabra Roth 1788. 
Monotropa hypophagos Monotropa hypophagos Dumortier, Bull. Soc. Roy. Bot. Belgique 4: 344. 1865.  
Monotropa hypophegea Monotropa hypophegea Wallroth, Sched. Crit. 191. 1822.BASIONYM: Monotropa hypopithys Linnaeus, var. glabra Roth 1788. 
Monotropa hypopithys forma cylindrica Monotropa hypopithys Linnaeus, forma cylindrica Andres, Verh. Naturhist. Vereines Preuss. Rheinl. Westphalens 66: 145. 1909.  
Monotropa hypopithys forma fusca Monotropa hypopithys Linnaeus, forma fusca Andres, Verh. Naturhist. Vereines Preuss. Rheinl. Westphalens 66: 145. 1909.  
Monotropa hypopithys forma glomerata Monotropa hypopithys Linnaeus, forma glomerata Andres, Verh. Naturhist. Vereines Preuss. Rheinl. Westphalens 66: 144. 1909.  
Monotropa hypopithys forma microcarpa Monotropa hypopithys Linnaeus, forma microcarpa Domin, Sitzungsber. Königl. Böhm. Ges. Wiss., Math.-Naturwiss. Cl. 1915: 13. 1915.  
Monotropa hypopithys forma minor Monotropa hypopithys Linnaeus, forma minor Domin, Sitzungsber. Königl. Böhm. Ges. Wiss., Math.-Naturwiss. Cl. 1915: 12. 1915.  
Monotropa hypopithys forma nutans Monotropa hypopithys Linnaeus, forma nutans Andres, Verh. Naturhist. Vereines Preuss. Rheinl. Westphalens 66: 145. 1909.  
Monotropa hypopithys forma pauciflora Monotropa hypopithys Linnaeus, forma pauciflora Haussknecht ex Andres, Verh. Bot. Vereins Prov. Brandenburg 54: 226. 1913.  
Monotropa hypopithys forma purpurascens Monotropa hypopithys Linnaeus, forma purpurascens Schutz ex Andres, Versamml. Bot.-Zool. Vereins Rheinl.-Westphalen 1912: 86. 1913.  
Monotropa hypopithys forma ramosa Monotropa hypopithys Linnaeus, forma ramosa (Rouy) Domin, Sitzungsber. Königl. Böhm. Ges. Wiss., Math.-Naturwiss. Cl. 1915: 19. 1915, non Andres 1909.BASIONYM: Hypopitys multiflora Scopoli, var. ramosa Rouy 1897. 
Monotropa hypopithys forma ramosa Monotropa hypopithys Linnaeus, forma ramosa Andres, Verh. Naturhist. Vereines Preuss. Rheinl. Westphalens 66: 144. 1909.  
Monotropa hypopithys forma rubra Monotropa hypopithys Linnaeus, forma rubra (Torrey) F. Seymour, Fl. New England, ed. 2. 427. 1982.BASIONYM: Monotropa lanuginosa Michaux, var. rubra Torrey 1846. 
Monotropa hypopithys forma subsessilis Monotropa hypopithys Linnaeus, forma subsessilis Domin, Sitzungsber. Königl. Böhm. Ges. Wiss., Math.-Naturwiss. Cl. 1915: 13. 1915.  
Monotropa hypopithys forma subuniflora Monotropa hypopithys Linnaeus, forma subuniflora Domin, Sitzungsber. Königl. Böhm. Ges. Wiss., Math.-Naturwiss. Cl. 1915: 18. 1915.  
Monotropa hypopithys forma vineosa Monotropa hypopithys Linnaeus, forma vineosa Andres, Verh. Naturhist. Vereines Preuss. Rheinl. Westphalens 66: 144. 1909.  
Monotropa hypopithys subvar. atricha Monotropa hypopithys Linnaeus, subvar. atricha Domin, Sitzungsber. Königl. Böhm. Ges. Wiss., Math.-Naturwiss. Cl. 1915: 17. 1915.  
Monotropa hypopithys subvar. piligera Monotropa hypopithys Linnaeus, subvar. piligera Domin, Sitzungsber. Königl. Böhm. Ges. Wiss., Math.-Naturwiss. Cl. 1915: 17. 1915.  
Monotropa hypopithys var. americana Monotropa hypopithys Linnaeus, var. americana (de Candolle) Domin, Sitzungsber. Königl. Böhm. Ges. Wiss., Math.-Naturwiss. Cl. 1915: 24. 1915.BASIONYM: Hypopitys multiflora Scopoli, var. americana de Candolle 1839. 
Monotropa hypopithys var. carnea Monotropa hypopithys Linnaeus, var. carnea Schutz ex G. Martens & Kemmler, Fl. Wuertemberg, ed. 3. 296. 1882.  
Monotropa hypopithys var. fimbriata Monotropa hypopithys Linnaeus, var. fimbriata (A. Gray) Domin, Sitzungsber. Königl. Böhm. Ges. Wiss., Math.-Naturwiss. Cl. 1915: 24. 1915.BASIONYM: Monotropa fimbriata A. Gray 1873. 
Monotropa hypopithys var. glaberrima Monotropa hypopithys Linnaeus, var. glaberrima H. Hara, J. Jap. Bot. 14: 427. 1938.  
Monotropa hypopithys var. glabra Monotropa hypopithys Linnaeus, var. glabra Roth, Tent. Fl. Germ. 1: 180. 1788. TYPE: GERMANY:
Monotropa hypopithys var. gracilescens Monotropa hypopithys Linnaeus, var. gracilescens Domin, Sitzungsber. Königl. Böhm. Ges. Wiss., Math.-Naturwiss. Cl. 1915: 21. 1915.  
Monotropa hypopithys var. hypophagos Monotropa hypopithys Linnaeus, var. hypophagos (Dumortier) Andres, Verh. Naturhist. Vereines Preuss. Rheinl. Westphalens 66: 145. 1909.BASIONYM: Monotropa hypophagos Dumortier 1865. 
Monotropa hypopithys var. hypophegea Monotropa hypopithys Linnaeus, var. hypophegea (Wallroth) Fries, Novit. Fl. Suec. Alt. 115. 1828, nom. illegit.BASIONYM: Monotropa hypophegea Wallroth 1822; Monotropa hypopithys Linnaeus, var. glabra Roth 1788. 
Monotropa hypopithys var. japonica Monotropa hypopithys Linnaeus, var. japonica (Franchet & Savatier) Franchet & Savatier, Enum. Pl. Jap. 2: 428. 1879.BASIONYM: Monotropa japonica Franchet & Savatier 1875. 
Monotropa hypopithys var. latisquama Monotropa hypopithys Linnaeus, var. latisquama (Rydberg) Kearney & Peebles, J. Wash. Acad. Sci. 29: 487. 1939.BASIONYM: Hypopitys latisquama Rydberg 1913. 
Monotropa hypopithys var. rubra Monotropa hypopithys Linnaeus, var. rubra (Torrey) Farwell, Amer. Midl. Naturalist 10: 39. 1926.BASIONYM: Monotropa lanuginosa Michaux, var. rubra Torrey 1846. 
Monotropa hypopithys var. tomentosa Monotropa hypopithys Linnaeus, var. tomentosa Velenovsky, Allg. Bot. Z. Syst. 10: 34. 1904.  
Monotropa hypoxya Monotropa hypoxya Sprengel, Syst. Veg. 2: 317. 1825, nom. illegit.BASIONYM: Monotropa hypophegea Wallroth 1822; Monotropa hypopithys Linnaeus, var. glabra Roth 1788. 
Monotropa japonica Monotropa japonica Franchet & Savatier, Enum. Pl. Jap. 1: 297. 1875. TYPE: JAPAN:
Monotropa laevipetala Monotropa laevipetala S. Gibson, Phytologist 1: 201. 1842.  
Monotropa lanuginosa var. glabriuscula Monotropa lanuginosa Michaux, var. glabriuscula Torrey, Fl. New York 1: 457. 1846. TYPE: NEW YORK: Yates Co.: Without data, Sartwell s.n. (holotype: ?).
Monotropa lanuginosa var. rubra Monotropa lanuginosa Michaux, var. rubra Torrey, Fl. New York 1: 457. 1846.  
Monotropa latisquama Monotropa latisquama (Rydberg) Hultén, Fl. Alaska 1216. 1948.BASIONYM: Hypopitys latisquama Rydberg 1913. 
Monotropa multiflora forma pauciflora Monotropa multiflora Fritsch, forma pauciflora (Haussknecht ex Andres) Margittai, Magyar Bot. Lapok 12: 244. 1913.BASIONYM: Monotropa hypopithys Linnaeus, forma pauciflora Haussknecht ex Andres 1913. 

Specimens

Country State County Date Specimen Notes Image
USA Florida Walton Co. 13 Oct 1994 A. Schotz 1190
USA Florida Okaloosa Co. 27 Jun 1994 A. Schotz 1100
USA Florida Marion Co. 10 Feb 1985 R. Lassiter 2193
USA Florida Lake Co. 11 Dec 1979 R. F. Daubenmire s.n.
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